The Bivalves portal

"Acephala", from Ernst Haeckel's Kunstformen der Natur (1904)

Bivalvia (/bˈvælviə/) or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed by a calcified exoskeleton consisting of a hinged pair of half-shells known as valves. As a group, bivalves have no head and lack some typical molluscan organs such as the radula and the odontophore. Their gills have evolved into ctenidia, specialised organs for feeding and breathing.

Common bivalves include clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. Majority of the class are benthic filter feeders that bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as scallops and file shells, can swim. Shipworms bore into wood, clay, or stone and live inside these substances.

The shell of a bivalve is composed of calcium carbonate, and consists of two, usually similar, parts called valves. These valves are for feeding and for disposal of waste. These are joined together along one edge (the hinge line) by a flexible ligament that, usually in conjunction with interlocking "teeth" on each of the valves, forms the hinge. This arrangement allows the shell to be opened and closed without the two halves detaching. The shell is typically bilaterally symmetrical, with the hinge lying in the sagittal plane. Adult shell sizes of bivalves vary from fractions of a millimetre to over a metre in length, but the majority of species do not exceed 10 cm (4 in).

Bivalves have long been a part of the diet of coastal and riparian human populations. Oysters were cultured in ponds by the Romans, and mariculture has more recently become an important source of bivalves for food. Modern knowledge of molluscan reproductive cycles has led to the development of hatcheries and new culture techniques. A better understanding of the potential hazards of eating raw or undercooked shellfish has led to improved storage and processing. Pearl oysters (the common name of two very different families in salt water and fresh water) are the most common source of natural pearls. The shells of bivalves are used in craftwork, and the manufacture of jewellery and buttons. Bivalves have also been used in the biocontrol of pollution.

Bivalves appear in the fossil record first in the early Cambrian more than 500 million years ago. The total number of known living species is about 9,200. These species are placed within 1,260 genera and 106 families. Marine bivalves (including brackish water and estuarine species) represent about 8,000 species, combined in four subclasses and 99 families with 1,100 genera. The largest recent marine families are the Veneridae, with more than 680 species and the Tellinidae and Lucinidae, each with over 500 species. The freshwater bivalves include seven families, the largest of which are the Unionidae, with about 700 species. (Full article...)

Selected article

Blood Ark Clam

The Blood ark clam is an ark clam with a fuzzy appearance, due to its periostracum. It is one of only a few molluscs that have red blood. (Read more…)

Selected biography

Constantine Samuel Rafinesque was a french polymath who studied botany, malacology, linguistics and anthropology. Educated in France, he immigrated to the United States at age of 19 where he befriended many botanists. At 21 he immigrated to Italy where he married. He would acquire 60,000 shells, which were lost after his ship sunk while immigrating back to the US. He was the first to use the term evolution, and would be honored by Charles Darwin. He however relied more on speculation than Darwin, for which he faced much criticism. (Read more...)

Did you know?

... that juvenile mangrove jingle clams can crawl up mangrove trees?

… that Spondylus are called spiny oysters, but possess eyes and are closer related to Scallops?

… that scaphopods are the closest relatives of bivalves?

In the news

5 July 2026 – Somalia–Turkey relations
The Puntland government issues a statement claiming exclusive control over its marine resources, citing its constitution and Somalia's federal framework to block SOMTURK, a joint venture between Somalia's fisheries ministry and Turkey's OYAK fund, from operating without approval. SOMTURK holds a federal licence for fishing permits across Somalia's economic zone. (Garowe Online) (Horseed Media) (Hiiraan Online)
20 May 2026 – Somalia–United States relations
Government of Puntland officials hosted a high-level United States delegation in the presidential palace in Garowe following the expiration of the mandate of Somalia’s federal government, where U.S. representatives sought to strengthen economic opportunities in Puntland’s oil exploration, fisheries, and mineral development sectors. (Garowe Online) (Horseed Media) (FTL Somalia)
16 May 2026 –
Solomon Islands reinstates a ban on dolphin exports two days after its repeal, citing concerns over the country's fisheries standards and tuna export industry. (AFP via The Manila Times) (RNZ)
12 February 2026 – China–Japan relations
The Japanese fisheries agency seize a Chinese fishing vessel and arrest its captain who failed to stop for inspection within Japan's exclusive economic zone off Nagasaki Prefecture, marking the first such seizure since 2022. (AFP via Philippine Daily Inquirer)
27 December 2025 – Lithium mining in Chile
Chilean state-owned mining company Codelco and private miner SQM create a joint venture to develop lithium in the Atacama salt flat until 2060, with the Chilean state set to receive the majority of operating margins. (AFP via Barron's)
8 December 2025 –
Singapore publishes the Albatross file, revealing previously-classified information about the Proclamation of Singapore and events leading up to the independence of Singapore for the first time in public view. (CNA)

Selected image

''Spondylus varius''
Spondylus varius


Spondylus varius, a large spiny oyster from off the coast of Africa.

Lists of Bivalves

Major Topics

  • Introductory articles
Bivalvia, Scallop, Mussel, clam, Freshwater bivalves, freshwater mussel,
  • Anatomy of hard parts
Bivalve shell, Adductor muscles (bivalve) Mantle (mollusc), periostracum, resilifer, resilium, Ligament (bivalve), Umbo (bivalve), Valve (mollusc),
  • Anatomy by systems
Digestive system of bivalves, respiratory system of bivalves, circulatory system of bivalves, excretory system of bivalves, sensory organs of bivalves, nervous system of bivalves, reproductive system of bivalves


  • Bivalves with significant positive human impact
As food Ornaments, pearls, etc. For other sciences
Scallop species
Oyster species
Venerid species
Quahog
Mussel species
Placopecten magellanicus
and many others
Nacre
Windowpane oyster
Quahog
Astarte (bivalve)
Giant clam
Jingle shell
Feather oysters
Chione species
Austrovenus species
Shelled taxa are valuable in archaeological and paleontological studies
  • Bivalves with significant negative human impact
Most invasive in freshwater Most invasive in saltwater Vectors for diseases
Zebra mussel
Corbicula fluminea
Mytella strigata
Limnoperna fortunei
Alathyria condola

Associated Wikimedia

The following Wikimedia Foundation sister projects provide more on this subject:

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