Bacteria secrete three different categories of cellulases including endoglucanases (EC 3.2.1.4), [beta]-glucosidases (EC 3.2.1.21) and cellobiohydrolase (EC 3.2.1.91) which act synergically to degrade complex cellulose material into simple sugar components (Matsui et al., 2013; Beeson et al., 2015).